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This wide-ranging course explains the key concepts of evolutionary science and investigates how these account for the characteristics of living organisms and the history of life on Earth. Using information from the living world and from fossil records, you'll learn how natural selection and other evolutionary processes produce changes in genes and populations over different timescales; how new species originate; and how large-scale evolutionary patterns are generated. Other topics include the reconstruction of evolutionary relationships and the ways in which humans influence the evolution of other species.

Modules count towards OU qualifications

OU qualifications are modular in structure; the credits from this undergraduate-level module could count towards a certificate of higher education, diploma of higher education, foundation degree or honours degree.

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Module code
Study level
3 10 6
Study method
Distance Learning
Module cost
See Module registration
Entry requirements
See Am I ready?

Student Reviews

“Everything about this course was brilliant. I thought the course textbook (Futuyma's 'Evolution'(FE)) was excellent and I constantly found myself...”
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“As several people have noted in their reviews, this is a very intensive course, but ultimately rewarding also. I personally...”
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What you will study

This course gives a broad treatment of both the biological and the palaeontological aspects of evolution, and so will be particularly rewarding to anyone who has an interest in life, earth or environmental sciences.

In particular, the course demonstrates how small genetic changes within populations (microevolution) can occur both through natural selection (including sexual selection) and as a consequence of other effects, and how such processes can lead to adaptive and other differences between populations. It considers how new species can arise; how evolutionary relationships are reconstructed; and how evolutionary changes in the genetic controls on development – and variation in the rates of speciation and of extinction – can lead to large-scale evolutionary patterns (macroevolution). It also looks at how humans have been (and are) affecting the evolution both of themselves and other contemporary organisms.

The course is based on the book Evolution by Douglas J. Futuyma (Sinauer, 2009), which is provided. A full colour Companion Text guides the student through the course books. It provides links to the additional components, including exercises and assignments designed to develop and test mastery of the learning outcomes of the course. Computer-based work involves interactive learning packages, video presentations, analytical software on DVD and a tutor-group wiki activity. You’ll also have access to a website and both tutor-group and course-wide online forums. For practical investigations, you’ll use a home kit of replica fossil specimens, supported by a Practical Booklet. The outcomes of these investigations are assessed in the tutor-marked assignments. Other supporting material includes an Introduction and Guide to the course (with included Glossary), and a Specimen Examination Paper.

Following introductions to the scientific basis of evolutionary theory and the key phenomena of adaptation and organised biological diversity, the methods for reconstructing evolutionary relationships are addressed with the aid of an interactive learning package and tree-building software. Patterns of evolution, as inferred from comparative studies, the fossil record, and the distributions of organisms are considered next. Supporting material includes a multimedia case study on the evolution of plumage in dinosaurs and birds (which is revisited in later parts of the course). There is also an exercise based on a set of plaster replicas of fossil shells supplied in the Home Kit.

The focus then moves on to the sources of genetic variation, how it affects the growth and form of organisms, and the investigation of genetic diversity within populations, all of which prepares you for the project work. Sexual selection and the issues of conflict and cooperation within species are explored next.

Then follow explorations of species concepts, how new species evolve, and life history evolution and its implications for the diversity of reproductive modes. These topics are supported by multimedia activities. The rapidly expanding field of developmental evolution is also considered, with a return to the plumage case study by way of illustration.

Finally, after considering some general broad issues, the course finishes with a look at the impact of humans on the evolution both of themselves and of other organisms.

Teaching and assessment

Support from your tutor

You will have a tutor who will help you with the study material and mark and comment on your written work, and who you can ask for advice and guidance. You will also be offered online tutorials, led by your tutor.

Contact our Student Registration & Enquiry Service if you want to know more about study with The Open University before you register.


The assessment details for this course can be found in the facts box above.

You can choose whether to submit your tutor-marked assignments (TMAs) on paper or online through the eTMA system. You may want to use the eTMA system for some of your assignments but submit on paper for others. This is entirely your choice.

Future availability

The details given here are for the course that starts in February 2014 when it will be available for the last time.


As a student of The Open University, you should be aware of the content of the Module Regulations and the Student Regulations which are available on our Essential documents website.

Course work includes:

4 Tutor-marked assignments (TMAs)
No residential school


This is a Level 3 course. Level 3 courses build on study skills and subject knowledge acquired from studies at Levels 1 and 2. They are intended only for students who have recent experience of higher education in a related subject, preferably with the OU. We advise against choosing S366 as your first Level 3 course.

To study this Level 3 course you need:

  • a background in science such as you would get from our Level 1 science course Exploring science (S104)
  • a pass in at least two Level 2 biology, earth science or environmental science courses such as Geology (S276) and Environmental science (S216) (or the equivalent)
  • to be competent at mathematics, to approximately the level of Maths for science (S151)
  • to be able to interpret graphs and tables of data
  • to be able to carry out some practical investigations involving measurement of specimens and analysis of the results.

Your regional or national centre will be able to tell you where you can see reference copies of the Level 1 and 2 courses. 

It is essential that you establish whether or not your background and experience give you a sound basis from which to tackle the course, since students who are appropriately prepared have the best chance of completing their studies successfully. The Science Faculty have produced a booklet, Are You Ready For S366? It will help you to decide whether you already have the recommended background knowledge or experience to start the course or whether you need a little extra preparation. This can be viewed or printed from the Are you ready for science? website. 

If you have any doubt about the suitability of the course, please contact our Student Registration & Enquiry Service.


Start End Fee
- - -

No current presentation - see Future availability

This module is expected to start for the last time in February 2014.

Ways to pay for this module

Open University Student Budget Account

The Open University Student Budget Accounts Ltd (OUSBA) offers a convenient 'pay as you go' option to pay your OU fees, which is a secure, quick and easy way to pay. Please note that The Open University works exclusively with OUSBA and is not able to offer you credit facilities from any other provider. All credit is subject to status and proof that you can afford the repayments.

You pay the OU through OUSBA in one of the following ways:

  • Register now, pay later - OUSBA pays your module fee direct to the OU. You then repay OUSBA interest-free and in full just before your module starts. 0% APR representative. This option could give you the extra time you may need to secure the funding to repay OUSBA.
  • Pay by instalments – OUSBA calculates your annual fees and spreads them out over up to a year, enabling you to pay your fees monthly and walk away with a qualification without any further debt. APR 5.1% representative.

Read more about Open University Student Budget Accounts (OUSBA).  

Employer sponsorship

Studying with The Open University can boost your employability. OU qualifications are recognised and respected by employers for their excellence and the commitment they take to achieve one. They also value the skills that students learn and can apply in the workplace.

More than one in 10 OU students are sponsored by their employer, and over 30,000 employers have used the OU to develop staff so far. If the qualification you’ve chosen is geared towards your job or developing your career, you could approach your employer to see if they will sponsor you by paying some or all of the fees. 

  • Your employer just needs to complete a simple form to confirm how much they will be paying and we will invoice them.
  • You won’t need to get your employer to complete the form until after you’ve chosen your modules.  

For more information about employer sponsorship speak to an adviser or request a call back.

Credit/debit card

You can pay part or all of your tuition fees upfront with a debit or credit card when you register for each module. 

We accept American Express, Maestro (UK only), Mastercard, Visa/Delta and Visa Electron. 

Gift vouchers

You can pay for part or all of your tuition fees with OU gift vouchers. Vouchers are currently available in the following denominations, £10, £20, £50 and £100. 

Mixed payments

We know that sometimes you may want to combine payment options. You may, for example wish to pay part of your tuition fee with a debit card and pay the remainder in instalments through an Open University Student Budget Accounts (OUSBA).

For more information about combining payment options, speak to an adviser or request a call back.

Note: Your permanent address/domicile will affect your fee status and therefore the fees you are charged and any financial support available to you. The fees and funding information provided here is based upon current details for  year 1 August 2014 to 31 July 2015.
This information was provided on 23/07/2014.

What's included

Two main texts, supplementary texts, DVD-ROM, a website and a home kit containing two real fossil specimens and a set of plaster replicas.

If you have a disability

In general, students with disabilities may find the practical work in the kit exercise and the project challenging. Those with impaired sight or manual dexterity may need an assistant. There is an option of a computer project for students who are unable to complete the other project options. The study materials are available in Adobe Portable Document Format (PDF). Components may not be fully accessible using a screen reader and mathematical, scientific, and foreign language materials may be particularly difficult to read in this way. Transcripts of video material will also be available. The books are available in a comb-bound format. Any queries on this subject should come to our Student Registration & Enquiry Service in the first instance.

If you have particular study requirements please tell us as soon as possible, as some of our support services may take several weeks to arrange. Find out more about our services for disabled students.